Friday 31 May 2013

Demetrias atricapillus (Linnaeus, 1758)

Regular viewers may have noticed by now quite how disproportionately species seem to group as described in 1758, with only one european spider (Enoplognatha ovata) pre-dating this (and even that's disputed).

This comes down to the history of taxonomy. Yes, you're about to get a recap.

In 1753, Carolus Linnaeus (spellings vary), a Swiss botanist, gave the Linnaean Binomial its public premiere in a book on plants. His systematic grouping of organisms based on relatedness (an early nod to evolution, as you'll note that the official birthday of creator-not-assumed evolution was the publication of On the Origin of Species by Means of Natural Selection in November 1859, over a century later) introduced a binomial naming system that paved the way for easy indexing of similar species and allowed naturalists to include either a descriptive identifier or a reference to something famous (or, in some cases, themselves). It was a more-or-less instant hit.

Five years later (in 1758), the botanist produced a zoological work, and the year of its publication was set as the official start of the attribution of binomial names to animals - that is to say, when taxonomists are trawling through records to try and find the first reference to an animal (and thereby credit a past naturalist with originally describing it), they don't have to bother looking at records older than that, which is handy because naming systems were more-or-less however the author pleased, and descriptions could be less than detailed.

However, as previously noted, the Linnaean binomial had been introduced - and quite popular - for plants five years previously, and quite a few naturalists had adopted it for other groups before Linnaeus himself applied it to everything in his Systema Naturae (that 1758 zoological work we mentioned earlier). Most of them are simply disregarded, but some - most notably Carl Alexander Clerck's(wiki link) 1757 treatise on spiders remain important works to this date, and so are less easily cast aside, which is why some spiders have a year-of-naming one year previous to the official start date for naming.

Okay, now that that's out of the way, on with the show...

Eukaryota
  Animalia
    Eumetazoa
      Bilateralia
        Nephrozoa
          Protostomia
            Ecdysozoa
              Arthropoda
                Hexapoda
                  Insecta
                    Dicondylia
                      Pterygota
                        Manopterygota
                          Neoptera
                            Eumetabola
                              Endopterygota
                                Coleopterida
                                  Coleoptera
                                    Adephaga
                                      Carabidae
                                        Harpalinae
                                          Lebiitae
                                            Lebiini
                                              Demetriadina

Note that outside of a few very well-studied groups of invertebrate, the use of supertribes (Lebiitae) and subtribes (Demetriadina) is extremely rare. Partially because evolution does not work at consistent speeds across the natural kingdom, but mostly because the more you find out about the relationships within a taxon, the more accurately you can tidy up their taxonomy. So while I would argue that ornithologists tend to oversplit (if an unbiased entomologist looked at a reed bunting and a sparrow, he or she would almost certainly keep them in one family, and quite possibly they wouldn't get anything more than a subgenus to separate them), entomologists tend to be working with taxa that have become so large that they are essentially meaningless, and so the finer splitting of higher taxonomy better represents the evolutionary history of a clade... 


Moving on - 

Demetrias atricapillus
(Linnaeus, 1758)

Demetrias atricapillus Bosham West Sussex UK
  This minute European ground beetle was photographed in West Sussex, UK in between a field and a saltmarsh... And, despite having been recognised as a separate species at least since Pope Benedict the 16th's clogs were popped, it has no common name.

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